Thousands of people died needlessly as a result of government mistakes in the handling of the coronavirus pandemic, Dominic Cummings has claimed.
The PM's former chief aide said Boris Johnson had initially dismissed Covid as a "scare story" and said the UK had been too slow to lock down.
And he claimed Health Secretary Matt Hancock had lied on multiple occasions and should have been fired.
"Tens of thousands of people died, who didn't need to die," said Mr Cummings.
In a marathon seven hour evidence session, Mr Cummings said: "The truth is that senior ministers, senior officials, senior advisers like me fell disastrously short of the standards that the public has a right to expect of its government in a crisis like this."
He added: "I would like to say to all the families of those who died unnecessarily how sorry I am for the mistakes that were made and for my own mistakes at that."
Mr Cummings - who was forced out of Number 10 at the end of last year after an internal power struggle - said those on the front line of the pandemic were like "lions" being "led by donkeys".
But he claimed Mr Johnson had told him he liked to be surrounded by "chaos" in Downing Street, because it meant everyone had to look to the PM "to see who is in charge".
Asked if he thought Mr Johnson was a "fit and proper person" to get the UK through the pandemic, Mr Cummings replied: "No."
Mr Johnson hit back at some of his former aide's allegations at Prime Minister's Questions, insisting that the government's priority had always been to "save lives".
But Mr Cummings said that despite nearly losing his own life to Covid, the PM thought the first national lockdown, on 23 March last year, had been a mistake - and he was against a so-called "circuit breaker" lockdown in autumn 2020 for economic reasons, despite scientific evidence it would have saved lives.
Asked on what basis the PM had taken this decision, Mr Cummings said: "He wasn't taking any advice, he was just taking his own decision that he was going to ignore the advice."
He added: "The cabinet wasn't involved or asked."
Asked if the prime minister had said he would rather see "bodies pile high" than take the country into a third lockdown, as reported by the BBC, Mr Cummins said: "I heard that in the prime minister's study."
Mr Johnson has denied making these comments.
Care homes
The former adviser reserved his most savage criticism for Matt Hancock, claiming the health secretary should have been fired for "15 to 20" different things - accusing him of "criminal, disgraceful behaviour that caused serious harm".
He said he had repeatedly called on Mr Johnson to sack Mr Hancock, calling him "completely incapable of doing the job".
The prime minister had been furious when he came back to work after recovering from coronavirus to find that untested patients had been discharged to care homes in England, thereby allowing the virus to spread, Mr Cummings claimed.
Both he and the PM had been told "categorically in March that people will be tested before they went back to care homes".
Subsequent government claims about putting a "shield" around care homes were "complete nonsense", he said.
Committee chairman Greg Clark said Mr Cummings must provide evidence to back up his claims about Mr Hancock.
Further claims
In other explosive claims, Mr Cummings said:
- Mr Johnson initially thought Covid-19 was just a "scare story" and the "new swine flu"
- The PM had offered to "get (Chief Medical Officer) Chris Whitty to inject me live on TV" to show there was nothing to fear from the virus
- Mr Johnson had never wanted tougher border controls to prevent the spread of the virus, as he wanted to be like the mayor in film Jaws, who kept beaches open despite the threat of a killer shark
Mr Cummings was questioned by the Health and Science select committees, for their inquiry into "lessons learnt" on the government's response to the pandemic.
Before the hearing began, Mr Cummings tweeted a picture of a whiteboard on which the government's "plan B" for the first wave of the virus was sketched out.
He told the MPs the original plan had been for limited intervention, with the hope of achieving "herd immunity", and he was "completely baffled" by Downing Street's denial of this.
He said the herd immunity strategy was abandoned when the scale of the death toll that would result from it became clear.
Barnard Castle
Mr Cummings was frank about some of his own failings - and admitted that his defence of a trip to County Durham during the first lockdown had "undermined public confidence".
But he said he had made the journey due to threats against his family - and he defended his claim that he had driven his family to Barnard Castle to test his eyesight, saying the story was too "weird" to have been made up.
He also gave his version of his departure from Downing Street, saying his relationship with Mr Johnson had broken down after the second national lockdown in October.
"The fact that his girlfriend (Carrie Symonds) also wanted rid of me was relevant but not the heart of the problem," he told MPs.
He said he regarded Mr Johnson as "unfit for the job" and was "trying to create a structure around him to try and stop what I thought were extremely bad decisions and push other things through against his wishes".
He also described how a meeting of the government's emergency Cobra committee on 12 March had been disrupted by US President Donald Trump wanting to start a "bombing campaign" against Iraq, Mr Cummings said.
'Save lives'
At the same time, he added, Mr Johnson's partner Carrie Symonds was "going crackers" about a "completely trivial" story in the Times regarding the dog she co-owned with the prime minister.
Labour leader Sir Keir Starmer quoted Mr Cummings at Prime Minister's Questions, as he called on the PM to apologise for his "complacency" at the start of the pandemic, which he said had led to "needless deaths".
He also asked Mr Johnson if - as Mr Cummings has claimed - he had delayed a lockdown in autumn last year because "Covid was only killing 80-year-olds". The PM did not respond to this allegation.
Mr Johnson told MPs: "The handling of this pandemic has been one of the most difficult things this country has had to do for a very long time and none of the decisions have been easy," he said.
"To go into a lockdown is a traumatic thing for a country, to deal with a pandemic on this scale has been appallingly difficult, and we have at every stage tried to minimise loss of life, to save lives, to protect the NHS and we have followed the best scientific advice that we can."
He added that he had not seen Mr Cummings' evidence to the select committee.
More than 127,000 people diagnosed with coronavirus have died in the UK since the start of the pandemic.
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